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David Davis
21262 Genoa Road
Linneus, MO 64653
Phone: 660 895-5121
FAX: 660 895=5122
Email:
DavisDK@missouri.edu
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September 25, 2000
SEASONAL NET ENERGY AVAILABILITY IN
ROTATIONALLY STOCKED PASTURES AT FOUR STOCKING RATES
Jim Gerrish
Research Assistant Professor,
University of Missouri - Forage Systems Research Center,
21262 Genoa Road, Linneus MO, 64653
(GerrishJ@missouri.edu)
Abstract
Nutrient intake by grazing animals is affected by both forage availability and
forage quality. Yearling steers were rotationally stocked from April to
September for four years at four stocking rates (300, 600, 900, 1200 pound
liveweight/acre) on cool-season grass-legume pastures to determine the effects
of stocking rate on forage availability and quality through the season. Lower
stocking rate resulted in higher forage availability but lower quality forage
while higher stocking rates produced less available forage but higher quality
forage. Peak forage availability at highest stocking rate occurred 58 days
earlier than for the lowest stocking rate. While forage availability declined
through the season, net energy content of the forage remained nearly constant
through the season. The combined effect of available forage and net energy
content resulted in significantly less net energy available per acre for the
1200 pound/acre stocking rate compared to all other stocking rates. These
results indicate that forage availability is more likely to limit steer
performance on pasture than is forage quality. Steer performance after mid
-summer was less than predicted based on available net energy per acre. Factors
other than forage availability or quality evidently limit steer performance
after mid-summer.
Introduction: Both forage availability and forage quality may limit animal
performance in grazing situations. In many research trials, only one of these
parameters may be measured. Failing to take both into account often leads to
over-prediction of expected animal performance or erroneous conclusions about
what factor is limiting animal performance. In other cases both are measured but
are not combined to provide a meaningful relationship to animal requirements and
performance. Knowing both forage production and nutritive value of the forage
allows the researcher or grazing manager to determine the nutrient supply and
demand balance.
Net energy demand of most classes of livestock under given production and
environmental scenarios has been determined. Net energy content of forage can be
calculated from forage analysis and forage availability or accumulation rate can
be measured. With this information, supply and demand relationships can be
determined and appropriate stock policy and stocking rate decisions can be made.
Our objective was to compare yearling steer net energy demand per acre with
forage net energy available per acre and determine what factors most likely
limit gain of grazing yearlings.
Materials and Methods: This project determined net energy accumulation rates for
cool-season grass-legume pastures rotationally stocked at four stocking rates.
Energy demand of yearling steers grazing these pastures was determined based on
net energy requirements for maintenance and growth.
Pastures consisted of endophyte-free tall fescue (Festuca arundinacea Schreb.),
orchardgrass (Dactylis glomerata L.), and Kentucky bluegrass (Poa pratensis L.)
overseeded with red clover (Trifolium pratense L.) and birdsfoot trefoil (Lotus
corniculatus L.) at the beginning of the study in 1995. Within each block,
pastures were randomly assigned a stocking rate and then split to either
continuous or rotational stocking treatments. The rotational grazing cells
consist of 12 equal sized paddocks. Target stocking rates were 300, 600, 900, or
1200 lb liveweight per acre at turn-out as yearling steers weighing
approximately 575 lb/head. Sixteen 10 acre pastures were used in the study to
provide two replications of each treatment in a randomized complete block design
with split plot assignment of treatments. Only rotational grazing data is
presented in this paper due to limited space.
The study was conducted from 1996 through 1999 with grazing beginning in early
to mid-April and ending around September 10. First grazing cycle consisted of
daily rotation through the 12 paddocks and subsequent cycles usually consisted
of 2-day grazing periods with 22 day rest periods. All stocking rates were
managed on the same rotation frequency.
In each rotationally grazed pasture, four paddocks were sampled in each grazing
cycle to determine forage availability. Nine 3.2 ft2 quadrats were clipped both
pre- and post-grazing in each paddocks. Bulk wet weight was measured and a 150 g
(+/-) subsample was oven dried to determine forage dry matter. Oven dried
samples were retained from each clipping for forage analysis. All forage samples
were analyzed by NIRS to determine crude protein, ADF, and NDF levels. Each year
approximately 100 samples were analyzed through standard wet chemistry
procedures to determine the same parameters and calibrate the NIRS data. Net
energy for maintenance (NEm) was calculated from ADF value using the
relationship:
Forage NEm = 1.04-(0.0104 X ADF)
Availability of NEm per acre was calculated as:
Forage NEm/acre = Herbage mass (lb/acre) X NEm (Mcal/lb)
Net energy demand for yearling steers was determined by the following equations:
NEm=.077 X ((liveweight/2.205).75
NEg=.0493 X ((liveweight/2.205).75) X ((ADG/2.205)1.097)
Animal demand per acre was calculated as:
Required NE/acre = NE/head X Head/acre
Results and Discussion: Forage availability was significantly affected by
stocking rate with date of peak forage availability also being determined by
stocking rate (Figure 1). Increased animal demand of higher stocking rates
resulted in lower mean forage availability throughout the season. While peak
forage availability occurred on July 29 for 300 pound/acre stocking rate, the
same event occurred on June 8 for 1200 pound/acre stocking rate. By the end of
the grazing season, forage availability on 1200SR was approximately 35% of
300SR.
The NEm content of forage samples was very good throughout the season with mean
levels remaining above .65 Mcal/pound for all treatments (Figure 2). This data
represents whole plant samples so animal selection would be expected to result
in even higher level of dietary energy intake. The highest stocking rate
produced the highest NEm forage while lowest stocking rate produced the lowest
NEm forage. Crude protein content, while not reported in this paper, responded
to stocking rate similarly.
These two pieces of data suggest that animal performance was more limited by
declining forage availability as compared to forage quality. The observed NEm
levels should be adequate to produce average daily gain in excess of 1.75
pounds/day if intake were not limited. However, by mid-July all treatments were
gaining less than 1.0 pound/day. It may be that reduction in ADG at this time
was due more to environmental stress than forage conditions. Nighttime low
temperatures were frequently above 80°F with humidity in excess of 70%. These
conditions are not conducive to steers achieving high pasture intake.
Net energy available per acre was greatest throughout the season for the lowest
stocking rate and least for the highest stocking rate (Figure 3). Available net
energy declined rapidly after mid-June for 1200SR. During a typical two-day
grazing period, NE consumption by the steers would exceed 50% of the available
NE. Forage utilization in excess of 50% will usually result in depressed animal
intake.
Daily net energy demand declined through the season for all treatments with more
rapid decline occurring as stocking rate increased (Figure 4). The decline was
due to decreased rate of gain as the season progressed. It is very difficult to
say whether gain decreased due to lack of energy intake or other environmental
factors. For all but the highest stocking rate, both forage availability and net
energy content were at levels which would not normally be expected to restrict
voluntary intake.
Steer average daily gain did not meet expectations based on available forage and
net energy content of the forage. It appears that factors other than the
measured forage parameters determine intake level and steer performance. High
nighttime temperatures and relative humidity may limit grazing time and total
daily intake. Maintaining high quality pastures with adequate availability will
help maintain summer rate of gain on yearling steers but other factors such as
proper environmental adaptation of livestock, parasite management, and
minimizing stresses may be equally important.

Figure 1. Forage availability declined through the season as stocking rate increased.

Figure 2. Higher stocking rates produced higher net energy forage with little variance through the seasons

Figure 3. Net energy available per acre varied through the season and was significantly affected by stocking rate.

Figure 4. Daily net energy demand per acre declined through the season at all stocking rates due to decling average daily gain.
SEASONAL NET ENERGY AVAILABILITY IN
ROTATIONALLY STOCKED PASTURES AT FOUR STOCKING RATES
Jim Gerrish
Research Assistant Professor,
University of Missouri - Forage Systems Research Center,
21262 Genoa Road, Linneus MO, 64653
(GerrishJ@missouri.edu)
Stocking rate refers to the number of animals or the total liveweight of animals
assigned to a grazing unit for and extended period of time. As stocking rate
increases, the animal demand for nutrients from every acre in the grazing unit
increases. Nutrient density is greater in less mature forage than in older plant
material. Increased grazing pressure causes the animals to consume more plant
material and stimulates regrowth, thus high grazing pressure increases nutrient
density of the forage while lower grazing pressure decreases forage quality but
leaves more forage available per acre. The challenge facing grazing managers is
whether a lesser amount of high quality forage or a greater amount of lower
quality forage is more conducive to animal growth. This study shows that even
though high grazing pressure produces the highest quality forage, the amount
available to the grazing animal is inadequate to meet their needs for both body
maintenance and growth. A moderate stocking rate produces the best combination
of forage quality and quantity.
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